The extreme, workerless inquilines.

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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Samstag 28. September 2019, 11:47

And here is the new list:

Literature list Teleutomyrmex T Off New 2.pdf
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Samstag 28. September 2019, 16:58

Those last three are included as:

Gösswald, K., 1954, “Unsere Ameisen. I. Teil.” Stuttgart, Franckh'sche Verlagshandlung, 88 pp.

Gösswald, K., 1955, “Unsere Ameisen. II. Teil.” Stuttgart, Franckh'sche Verlagshandlung, 80 pp.

Gösswald, K., 1985, “Organisation und Leben der Ameisen.” Stuttgart, Wissenschaftliche Verlagsgesellschaft mbH, 355 pp.

Total: 90 references.
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Dienstag 1. Oktober 2019, 15:37

An official literature list of Teleutomyrmex Kutter, 1950.

Baroni Urbani, C., 1967, “Le distribuzioni geografiche discontinue dei Formicidi mirmecobiotici.” Archivio Botanico e Biogeografico Italiano, vol. 43 [=(4)12]: p. 355-365.

Baur, A., Buschinger, A., Zimmermann, F. K., 1993, “Molecular cloning and sequencing of 18S rDNA gene fragments from six different ant species.” Insectes Sociaux, vol. 40, p. 325-335.

Baur, A., Chalwatzis, N., Buschinger, A., Zimmermann, F. K., 1995, “Mitochondrial DNA sequences reveal close relationships between social parasitic ants and their host species.” Current Genetics, vol. 28, p. 242-247.

Baur, A., Sanetra, M., Chalwatzis, N., Buschinger, A., Zimmermann, F. K., 1996, “Sequence comparisons of the internal transcribed spacer region of ribosomal genes support close relationships between parasitic ants and their respective host species (Hymenoptera: Formicidae).” Insectes Sociaux, vol. 43, p. 53-67.

Bernard, F., 1967 ["1968"], “Faune de l'Europe et du Bassin Méditerranéen. 3. Les fourmis (Hymenoptera Formicidae) d'Europe occidentale et septentrionale.” Paris, Masson, 411 pp.

Blaimer, B. B., Ward, P. S., Schultz, T. R., Fisher, B. L., Brady, S. G., 2018, “Paleotropical Diversification Dominates the Evolution of the Hyperdiverse Ant Tribe Crematogastrini (Hymenoptera: Formicidae).” Insect Systematics and Diversity, vol. 2, no. 5, art. 3, p. 1-14.

Bolton, B., 1976, “The ant tribe Tetramoriini (Hymenoptera: Formicidae). Constituent genera, review of smaller genera and revision of Triglyphothrix Forel.” Bulletin of the British Museum (Natural History) (Entomology), vol. 34, p. 281-379.

Bolton, B., 1994, “Identification guide to the ant genera of the world.” Cambridge, Mass., Harvard University Press, 222 pp.

Bolton, B., 1995, “A taxonomic and zoogeographical census of the extant ant taxa (Hymenoptera: Formicidae).” Journal of Natural History, vol. 29, p. 1037-1056.

Bolton, B., 1995, “A new general catalogue of the ants of the world.” Cambridge, Mass., Harvard University Press, 504 pp.

Bolton, B., 2003, “Synopsis and Classification of Formicidae.” Memoirs of the American Entomological Institute, vol. 71, 370 pp.

Borowiec, L., 2014, “Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae).” Genus (Wroclaw), vol. 25, no. 1-2, p. 1-340.

Brown, W. L., Jr., 1973, “A comparison of the Hylean and Congo-West African rain forest ant faunas.” P. 161-185 in: Meggers, B. J., Ayensu, E. S., Duckworth, W. D. (ed.), 1973, “Tropical forest ecosystems in Africa and South America: a comparative review.” Smithsonian Institution Press, Washington, DC., viii + 350 pp.

Brun, R., 1952, “Das zentralnervensystem von Teleutomyrmex schneideri Kutt. (Hym. Formicid.). III. Mitteilung.” Mitteilungen der Schweizerischen Entomologischen Gessellschaft, vol. 25, no. 2, p. 73-86.

Brun, R., 1959, “Le cerveau des fourmis et des insectes en général comme instrument de formation des réflexes conditionnés.” P. 11-25 in: International Union of Biological Sciences, Union Internationale des Sciences Biologiques, Experimental Psychology and Animal Behaviour Section, 1959, “Animal psychology seminars: Strasbourg University, October 1956, and Brussels University, August 1957.” London, Pergamon Press, 148 pp.

Buschinger, A., 1970, “Neue Vorstellungen zur Evolution des Sozialparasitismus und der Dulosis bei Ameisen (Hym., Formicidae).” Biologisches Zentralblatt, vol. 88, no. 3, p. 273-299.

Buschinger, A., 1971, “Zur Verbreitung und Lebensweise sozialparasitischer Ameisen des Schweizer Wallis (Hym., Formicidae).” Zoologischer Anzeiger, vol. 186, no. 1/2, p. 47-59.

Buschinger, A., 1974, “Monogynie und Polygynie in Insektensozietäten.” P. 862-896 in: Schmidt, G. H. (ed.), 1974, “Sozialpolymorphismus bei Insekten. Probleme der Kastenbildung im Tierreich.” Stuttgart, Wissenschaftliche Verlagsgesellschaft mbH, xxiv + 974 pp.

Buschinger, A., 1974, “Polymorphismus und Polyethismus sozialparasitischer Hymenopteren.” P. 897-934 in: Schmidt, G. H. (ed.), 1974, “Sozialpolymorphismus bei Insekten. Probleme der Kastenbildung im Tierreich.” Stuttgart, Wissenschaftliche Verlagsgesellschaft mbH, xxiv + 974 pp.

Buschinger, A., 1985, “New records of rare parasitic ants (Hym., Form.) in the French Alps.” Insectes Sociaux, vol. 32, no. 3, p. 321-324.

Buschinger, A., 1986, “Evolution of social parasitism in ants.” Trends in Ecology and Evolution, vol. 1, p. 155-160.

Buschinger, A., 1987, “Teleutomyrmex schneideri Kutter, 1950 and other parasitic ants found in the Pyrenees (Hymenoptera, Formicidae).” Spixiana, vol. 10, no. 1, p. 81-83.

Buschinger, A., 1990, “Evolutionary transitions between types of social parasitism in ants, hypotheses and evidence.” P. 145-146 in: Veeresh, G. K., Mallik, B., Viraktamath, C. A. (eds.), 1990, “Social insects and the environment. Proceedings of the 11th International Congress of IUSSI, 1990.” New Delhi, Oxford & IBH Publishing Co., xxxi + 765 pp.

Buschinger, A., 1990, “Sympatric speciation and radiative evolution of socially parasitic ants. - Heretic hypotheses and their factual background.” Zeitschrift für Zoologische Systematik und Evolutionsforschung, vol. 28, p. 241-260.

Buschinger, A., 1995, “Nicht am Ende: Die “Endameise” Teleutomyrmex schneideri.” Ameisenschutz aktuell, vol. 9, p. 1-7.

Buschinger, A., 1999, “Wiederfund der sozialparasitischen Ameise Teleutomyrmex schneideri in der Schweiz.” Mitteilungen der Schweizerischen Entomologischen Gesellschaft, vol. 72, p. 277-279.

Buschinger, A., 2000, “Die “Endameise” Teleutomyrmex schneideri in der Schweiz: Erster Wiederfund nach 50 Jahren.” Ameisenschutz aktuell, vol. 14, p. 43-45.

Buschinger, A., 2009, “Social parasitism among ants: a review (Hymenoptera: Formicidae).” Myrmecological News, vol. 12, p. 219-235.

Casevitz-Weulersse, J., Galkowski, C., 2009, “Liste actualisée des fourmis de France (Hymenoptera, Formicidae).” Bulletin de la Société Entomologique de France, vol. 114, no. 4, p. 475-510.

Collingwood, C. A., 1956, “Ant hunting in France.” Entomologist, vol. 89, p. 105-108.

Collingwood, C. A., 1956, “A rare parasitic ant (Hym., Formicidae) in France.” Entomologist's Monthly Magazine, vol. 92, p. 197.

Csősz, S., Radchenko, A., Schulz, A., 2007, “Taxonomic revision of the Palaearctic Tetramorium chefketi species complex (Hymenoptera: Formicidae).” Zootaxa 1405, p. 1-38.

Cuesta, D., García, F., García-Tejero, S. & Espadaler, X., 2009, “Resumen charla: Aportaciones a la biología de Teleutomyrmex schneideri Kutter, 1950: primer caso de cría en cautividad. [Contributions to the biology of Teleutomyrmex schneideri Kutter, 1950: first case of keeping in captivity.]” [abstract]. Iberomyrmex, vol. 1, p. 24.

Cuesta-Segura, A.D., Espadaler, X., Garcia Garcia, F., 2017, “Hormigas de los brezales de Calluna cantábricos (NO España) (Hymenoptera: Formicidae). [Ants of the Cantabrian Calluna-heathlands (NW Spain) (Hymenoptera: Formicidae).]” Iberomyrmex, vol. 9, p. 25-43.

Cuesta-Segura, A. D., García García, F., Catarineu, C., García-Tejero, S., Espadaler, X., 2018, “Actualización de la Distribución y Hospedadores de la Hormiga Parasita Teleutomyrmex schneideri Kutter, 1950 en la Peninsula Ibérica (Hymenoptera: Formicidae).” Boletín de la Sociedad Entomológica Aragonesa (S.E.A.), nº 63 (31/12/2018), p. 235–239.

Dlussky, G. M., Fedoseeva, E. B., 1988 (“1987”), “[The origin and early stages of evolution of ants (Hymenoptera: Formicidae).]” P. 70-144 in: Ponomarenko, A. G. (ed.), “[The Creaceous biocenotic crisis and the evolution of insects.]” [In Russian.]. Nauka, Moscow. 231 pp.

Dlussky, G. M., Soyunov, O. S., Zabelin, S. I., 1990 [“1989”], “Muravji Turkmenistana.”, or “[Ants of Turkmenistan.]” [In Russian.]. Ashkabad, Ylym Press, 275 pp.

Espadaler, X., Cuesta, D., 2006, “Notas / Notes. Teleutomyrmex schneideri Kutter, 1950 en España (Hymenoptera, Formicidae).” Graellsia, vol. 62, nr. 2, p. 261-262.

Espadaker, X., Gómez, K., 2014, “Tetramorium biskrense Forel, 1904 en España y Portugal peninsulares (Hymenoptera, Formicidae).” Boletín de la Sociedad entomológica Aragonesa (SEA), vol. 55, p. 303-305.

García, F., 2019, “Parásitos y myrmecófilos. Hormigas parásitas sociales ibéricas.” LaMarabunta digital, Nueva Edición, Número 4, Junio 2019: p. 27-44.

Gösswald, K., 1952, “Zur Biologie und Histologie parasitär degenerierter Ameisenarten mit besonderer Berücksichtigung von Teleutomyrmex schneideri Kutter (Tribus Tetramorini).” Transactions of the IX-th International Congress of Entomology, Amsterdam, August 17-24 (1951), vol. 1, p. 446-448.
- His name is spelled “Göszwald” in this publication!

Gösswald, K., 1953, “Histologische Untersuchungen an der arbeiterlosen Ameise Teleutomyrmex schneideri Kutter (Hym. Formicidae).” Mitteilungen der Schweizerischen Entomologischen Gessellschaft, vol. 26, no. 2, p. 81-128.

Gösswald, K., 1954, “Unsere Ameisen. I. Teil.” Stuttgart, Franckh'sche Verlagshandlung, 88 pp.

Gösswald, K., 1955, “Unsere Ameisen. II. Teil.” Stuttgart, Franckh'sche Verlagshandlung, 80 pp.

Gösswald, K., 1985, “Organisation und Leben der Ameisen.” Stuttgart, Wissenschaftliche Verlagsgesellschaft mbH, 355 pp.

Hölldobler, B. K., & Wilson, E. O., 1990, “The Ants.” Cambridge, Mass., Belknap Press of Harvard University Press, xiv + 732 pp.

Hölldobler, B. K., & Wilson, E. O., 1998, “Journey to the Ants: A Story of Scientific Exploration.” Cambridge, Mass., Belknap Press of Harvard University Press, 304 pp.

Kiran, K., Karaman, C., Lapeva-Gjonova, A. & Aksoy, V., 2017, “Two new species of the “ultimate“ parasitic ant genus Teleutomyrmex Kutter, 1950 (Hymenoptera: Formicidae) from the Western Palaearctic.“ Myrmecological News, vol. 25, p. 145-155.

Kusnezov, N., 1954, “Phyletische Bedeutung der Maxillar- und Labialtaster der Ameisen.” Zoologischer Anzeiger, vol. 153, no. 1/2, p. 28-38.

Kutter, H., 1950, “Über eine neue, extrem parasitische Ameise. 1. Mitteilung.” Mitteilungen der Schweizerischen Entomologischen Gessellschaft, vol. 23, no. 2, p. 81-94.

Kutter, H., 1950, “Über zwei neue Ameisen.” Mitteilungen der Schweizerischen Entomologischen Gessellschaft, vol. 23, no. 3, p. 337-346.

Kutter, H., 1950, “Über Doronomyrmex und verwandte Ameisen. 2. Mitteilung.” Mitteilungen der Schweizerischen Entomologischen Gessellschaft, vol. 23, no. 3, p. 347-353.

Kutter, H., 1951, “Von dufttäuschenden Mörderinnen und berittener Königin.” Du: kulturelle Monatsschrift, vol. 11, no. 4, 44-47.

Kutter, H., 1952, “Über Plagiolepis xene Stärcke (Hym. Formicid.).” Mitteilungen der Schweizerischen Entomologischen Gessellschaft, vol. 25, no. 2, p. 57-72.

Kutter, H., 1963, “Miscellanea myrmecologica I.” Mitteilungen der Schweizerischen Entomologischen Gessellschaft, vol. 36, no. 1 and 2, p. 129-137.

Kutter, H., 1968 ("1967"), “Liste sozialparasitischer Ameisen.” Archives. Institut Grand-Ducal de Luxembourg (n.s.), vol. 33, p. 201-210.

Kutter, H., 1968 (“1969”), “Die sozialparasitischen Ameisen der Schweiz.” Neujahrsblatt herausgegeben von der Naturforschenden Gesellschaft in Zürich auf das Jahr 1969, vol. 171, (Ausgegeben am 31. Dezember 1968), p. 1-62.

Kutter, H., 1977, “Hymenoptera, Formicidae.” Insecta Helvetica. Fauna, vol. 6, p. 1-298.

Kutter, H., 1978, “Hymenoptera, Formicidae.” Insecta Helvetica. Fauna, vol. 6a, unpaginated, illustr.

Meyer, G., 1951, “Histologische Untersuchungen an einer arbeiterlosen Ameise Anergates atratulus Schenck.” Inaugural-Dissertation, Institut für Angewandte Zoologie, Würzburg.

Meyer, G. F., 1955, “Untersuchungen an einer parasitischen Ameise (Anergates atratulus Schenck).” Insectes Sociaux, vol. 2, no. 2, p. 163-171.

Reyes López, J., Martínez, A. B., 2011, “Notas Científicas: Nueva cita de Teleutomyrmex kutteri Tinaut, 1990 (Hym., Formicidae) para la Península Ibérica. New record of Teleutomyrmex kutteri Tinaut, 1990 (Hym., Formicidae) from the Iberian Peninsula.” Boletín de la Sociedad Entomológica Aragonesa (S.E.A.), nº. 49, p. 206.

Sanetra, M., 1996, “Systematik, Biologie und Phylogenie westpaläarktischer Ameisen der Tribus Tetramoriini unter besonderer Berücksichtigung molekularer Merkmale (Hymenoptera: Formicidae).” Thesis, Technische Hochschule Darmstadt, p, 1-229.

Sanetra, M., Buschinger, A., 2000, “Phylogenetic relationships among social parasites and their hosts in the ant tribe Tetramoriini (Hymenoptera: Formicidae).” European Journal of Entomology, vol. 97, no. 1, p. 95-117.

Sanetra, M., Felger, S., Buschinger, A., Zimmermann, F. K., 1998, “On the evolutionary history of social parasites in the ant tribe Tetramoriini (Hymenoptera: Formicidae).” P. 413 in: Schwarz, M. P., Hogendoorn, K. (eds.), 1998, “Social insects at the turn of the millennium. Proceedings of the XIII International Congress of IUSSI. Adelaide, Australia. 29 December 1998 - 3 January 1999.” Adelaide, XIII Congress of IUSSI, 535 pp.

Sanetra, M., Heinze, J., Buschinger, A., 1994, “Enzyme polymorphism in the ant genus Tetramorium Mayr and its social parasites (Hymenoptera: Formicidae).” Biochemical Systematics and Ecology, vol. 22, p. 753-759.

Schlick-Steiner, B. C., Steiner, F. M., Moder, K., Seifert, B., Sanetra, M., Dyreson, E., Stauffer, C., Christian, E., 2005, “A multidisciplinary approach reveals cryptic diversity in Western Palaearctic Tetramorium ants (Hymenoptera: Formicidae).” Molecular Phylogenetics and Evolution, vol. 40, p. 259-273.

Schönitzer, K., Lawitzky, G., 1987, “A phylogenetic study of the antenna cleaner in Formicidae, Mutillidae, and Tiphiidae (Insecta, Hymenoptera).” Zoomorphology (Berlin), vol 107, no. 5, p. 273-285.

Seifert, B., 1996, “Ameisen: beobachten, bestimmen.” Naturbuch Verlag, Augsburg, 351 pp.

Seifert, B., 2007, “Die Ameisen Mittel- und Nordeuropas.” Lutra Verlags- und Vertriebsgesellschaft, Tauer/Klitten, 368 pp.

Seifert, B., 2018, “The ants of Central and North Europe.” Lutra Verlags- und Vertriebsgesellschaft, Tauer, 407 pp.

Seifert, B., Buschinger, A., Aldawood, A., Antonova, V., Bharti, H., Borowiec, L., Dekoninck, W., Dubovikoff, D., Espadaler, X., Flegr, J. , Georgiadis, C., Heinze, J., Neumeyer, R., Ødegaard, F., Oettler, J., Radchenko, A., Schultz, R., Sharaf, M., Trager, J., Vesnić, A., Wiezik, M., Zettel, H., 2016, “Banning paraphylies and executing Linnaean taxonomy is discordant and reduces the evolutionary and semantic information content of biological nomenclature.” Insectes Sociaux, vol. 63, no. 2, p. 237-242.

Social Insects Specialist Group, 1996a, “Teleutomyrmex kutteri. The IUCN Red List of Threatened Species 1996: e.T21570A 9301532. http://dx.doi.org/10.2305/IUCN.UK.1996.RLTS.T 21570A9301532.en.”

Sociak Insects Specialist Group, 1996b, “Teleutomyrmex schneideri. The IUCN Red List of Threatened Species 1996: e.T21569A9301467. http://dx.doi.org/10.2305/IUCN.UK.19 96.RLTS.T21569A9301467.en.”

Steiner, F. M., Seifert, B. , Moder, K., Schlick-Steiner, B. C., 2010, “A multisource solution for a complex problem in biodiversity research: Description of the cryptic ant species Tetramorium alpestre sp.n. (Hymenoptera: Formicidae).” Zoologischer Anzeiger, vol. 249, p. 223-254.

Stumper, R., 1951, “Teleutomyrmex schneideri Kutter (Hym. Formicid.). II. Mitteilung. Über die Lebensweise der neuen Schmarotzerameise.” Mitteilungen der Schweizerischen Entomologischen Gessellschaft, vol. 24, no. 2, p. 129-152.

Stumper, R., 1956, “Etudes myrmécologiques LXXVII. Les sécrétions attractives des reines de fourmis.” Mitteilungen der Schweizerischen Entomologischen Gessellschaft, vol. 29, no. 4, p.373-380.

Stumper, R., Kutter, H., 1950, “Sur le stade ultime du parasitisme social chez les fourmis, atteint par Teleutomyrmex Schneideri (subtrib. nov.; gen. nov.; spec. nov. Kutter).” Comptes Rendus (Hebdomadaires) des Séances de l'Academie des Sciences, vol. 231, p. 876-878.

Tinaut, A., 1981, “Estudio de los Formícidos de Sierra Nevada.” Tesis Doctoral, Universidad de Granada, 463 pp.

Tinaut, A., 1990, “Teleutomyrmex kutteri, spec. nov. A new species from Sierra Nevada (Granada, Spain).” Spixiana, vol. 13, no. 2, p. 201-208.

Tinaut, A., Ruano, F., Martínez, M. D., 2005, “Biology, Distribution and Taxonomic Status of the Parasitic Ants of the Iberian Peninsula (Hymenoptera: Formicidae, Myrmicinae).” Sociobiology, vol. 46, no. 3, p. 449-489.

Wagner, H. C., Arthofer, W., Seifert, B., Muster, C., Steiner, F. M., Schlick-Steiner, B. C., 2017, “Light at the end of the tunnel: Integrative taxonomy delimits cryptic species in the Tetramorium caespitum complex (Hymenoptera: Formicidae).” Myrmecological News, vol. 25, p. 95–129.

Ward, P. S., Brady, S. G., Fisher, B. L., Schultz, T. R., 2015 (“2014”), “The evolution of myrmicine ants: Phylogeny and biogeography of a hyperdiverse ant clade (Hymenoptera: Formicidae).” Systematic Entomology, vol. 40, no. 1, p. 61-81. (Article first published online: 23 July 2014).

Ward, P. S., Brady, S. G., Fisher, B. L., Schultz, T. R., 2016, “Phylogenetic classifications are informative, stable, and pragmatic: the case for monophyletic taxa.” Insectes Sociaux, vol. 63, no. 4, p. 489-492.

Wegnez, P., Ignace, D., Lommelen, E., Hardy, M., Bogaert, J., Nilsson, C., 2015, “Redécouverte de Teleutomyrmex schneideri Kutter, 1950 dans les Alpes françaises (Hymenoptera: Formicidae).” Bulletin de la Société royale belge d’Entomologie/Bulletin van de Koninklijke Belgische Vereniging voor Entomologie, vol. 151, p. 52-57.

Wheeler, G. C., Wheeler, J., 1985, “A simplified conspectus of the Formicidae.” Transactions of the American Entomological Society, vol. 111, p. 255-264.

Wilson, E. O., 1963, “The Social Biology of Ants.” Annual Review of Entomology, vol. 8, no. 1, 345-368.

Wilson, E. O., 1963, “Social modifications related to rareness in ant species.” Evolution, vol. 17, p. 249-253.

Wilson, E. O., 1971, “The insect societies.” Cambridge, Mass., Harvard University Press, x + 548 pp.

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Re: The extreme, workerless inquilines.

Beitragvon Merkur » Dienstag 1. Oktober 2019, 19:50

Ad Baroni Urbani 1967!

Hi Teleutotje,
A couple of weeks ago you asked for information on a paper of Baroni Urbani 1967, here
I found out that I possess a reprint!
Here are the importent pages (I can send you a scan of the whole paper, if you wish).
Baroni-1-219.jpg
Deckblatt
Baroni-2-220.jpg
Titelblatt
Baroni-3-221.jpg
"Epimyrma"
Baroni-4-223.jpg
Page with Teleutomyrmex
Baroni-5-222.jpg
Summary

Die Arbeit ist ziemlich überholt, da man z. B. annahm, dass Anergates atratulus in Europa und Nordamerika konvergent entstanden sei. Heute sind wir sicher, dass diese extrem parasitisch abgewandelte Art zusammen mit ihren Tetramorium-Wirten aus Europa nach Nordamerika gebracht worden ist.
Die in der Arbeit ebenfalls erwähnten, ehemals für Sozialparasiten gehaltenen Arten der Gattung Xenometra wurden inzwischen als ergatoide Männchen-Form in der Gattung Cardiocondyla entlarvt, u. a. durch Baroni Urbani: Baroni Urbani, C. 1973a. Die Gattung Xenometra, ein objektives Synonym (Hymenoptera, Formicidae). Mitt. Schweiz. Entomol. Ges. 46: 199-201 (page 199, Xenometra as junior synonym of Cardiocondyla) - https://www.e-periodica.ch/digbib/view? ... :1973:46#5
Da inzwischen mehrere Gattungen einst des Sozialparasitismus „verdächtigter“ Arten sich als abweichend gestaltete Formen (meist Gynen) ganz normaler Ameisen erwiesen haben (z. B. Epixenus und Xenhyboma), darf man sich keinesfalls verwirren lassen durch die jüngeren Synonymisierungen von Ward et al.2015: Die Arten der Gattungen Anergates, Teleutomyrmex, Chalepoxenus und Myrmoxenus sind und bleiben echte Sozialparasiten, auch wenn jene Autoren sie mit ihren jeweiligen Wirtsgattungen synonymisiert haben!- Nicht alles ist gut, was aus den USA kommt, leider auch in der Myrmekologie. :(

MfG,
Merkur
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Zuletzt geändert von Merkur am Mittwoch 2. Oktober 2019, 08:52, insgesamt 1-mal geändert.
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Dienstag 1. Oktober 2019, 21:30

Dear Prof.,

I would like that very great!

Hope to see it soon….

Yours,

Teleutotje
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Samstag 19. Oktober 2019, 21:53

The fig. from Dlussky, G. M., Soyunov, O. S., Zabelin, S. I., 1990 [“1989”], “Muravji Turkmenistana.”, or “[Ants of Turkmenistan.]” [In Russian.]. Ashkabad, Ylym Press, 275 pp.:

http://antclub.ru/f/769/r22.gif
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Samstag 19. Oktober 2019, 21:57

In that fig. you have on top a Teleutomyrmex schneideri queen, at the bottem you have a Strongylognathus testaceus worker.
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Samstag 19. Oktober 2019, 22:18

And the text in that book about Teleutomyrmex:

Text.jpg
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Samstag 11. Januar 2020, 20:58

How F. Bernard speaks about Teleutomyrmex and Anergates:
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Tel 01.jpg
Tel 02.jpg
Tel 03.jpg
Tel 04.jpg
Tel 05.jpg
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Samstag 11. Januar 2020, 21:05

And B. Seifert in his last book:
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Dienstag 14. Januar 2020, 09:48

One small remark: I have my Teleutomyrmex-specimens, 1 male, 2 females, from October, 4, 2002.
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Mittwoch 22. Januar 2020, 20:58

Now, a new contribution to Myrmecological News. I'm still reading it but it looks great.

de la Mora, A., Sankovitz, M. & Purcell, J., 2020, "Ants (Hymenoptera: Formicidae) as host and intruder: recent advances and future directions in the study of exploitative strategies." Myrmecological News, Vol. 30, 53-71.

https://myrmecologicalnews.org/cms/inde ... Itemid=435
or
https://myrmecologicalnews.org/cms/inde ... format=raw and https://myrmecologicalnews.org/cms/inde ... format=raw
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Donnerstag 23. Januar 2020, 23:10

And

Stuart, R.J., 2002, “The behavioural ecology of social parasitism in ants.” P. 315-336 in: Lewis, E. E., Campbell, J. F., Sukhdeo, M. V. K. (eds), 2002, “The Behavioural Ecology of Parasites.” CABI Publishing, Wallingford, UK and New York, USA, 369 pp.

Don't know anything about this…

92 references.
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Re: The extreme, workerless inquilines.

Beitragvon Merkur » Freitag 24. Januar 2020, 16:14

@ Teleutotje: Diesen Buchbeitrag von R. J. Stuart 2002 kenne ich selbst nicht, und deswegen ein allgemeines Buch über Parasitismus für 120 $ zu kaufen, ist mir zuviel. Ich erwarte darin allerdings keine „großen“ Neuigkeiten. Robin hat selbst bei T. M. Alloway über sklavenhaltende Ameisen gearbeitet (zusammen haben wir u.a. in Québéc Harpagoxenus canadensis gesammelt). Üblicherweise hat man damals noch Sonderdrucke seiner Publikationen ausgetauscht, später dann per E-Mail PDFs, aber das kommt zunehmend aus der Mode…. :roll:
Gewöhnlich wird man als Spezialist für bestimmte Themen von Buch-Herausgebern bzw. Verlagen aufgefordert/gebeten, solche Beiträge zu liefern. Zunächst fühlt man sich geehrt. ;) Aber meist ist eine bestimmter Umfang vorgegeben. Da ist es oft aufwendig, viel Literatur zum Thema durchzuarbeiten, und natürlich möchte man auch eigene Ergebnisse unterbringen, wofür dann doch nur wenig Raum ist. Inzwischen lehne ich solche Anfragen einfach ab.
Der Beitrag von R. Stuart im MN-Artikel wird S. 58 zitiert mit:
„Likewise, Tetramorium inquilinum (=Teleutomyrmex schneideri) queens are small compared with their hosts, have unusually large tarsal claws, and spend much of their time riding around on the backs of their host queen (Stuart 2002).“ - Viel mehr dürfte in dem Buchbeitrag 2002 auch nicht stehen. ;)

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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Donnerstag 30. Januar 2020, 22:00

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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Samstag 14. März 2020, 16:32

Teleutomyrmex.

From AmeisenWiki, on 14/03/2020, from 12:30 till 12:35:

“Teleutomyrmex.
Teleutomyrmex ist eine Gattung aus der Unterfamilie Myrmicinae. Sie umfasst nur zwei Arten, T. schneideri (Westalpen, Pyrenäen, Kasachstan) und T. kutteri (Südspanien, Sierra Nevada, bisher nur ein Volk gefunden).
Es handelt sich um arbeiterinnenlose Sozialparasiten in Nestern von Tetramorium cf. impurum. Die Tiere sind extrem selten. Auch von T. schneideri wurden seit der Entdeckung (1949 im Schweizer Wallis) bis 2006 nicht mehr als 15 Völker gefunden.
Die stark physogastrischen Königinnen hocken fast permanent auf dem Rücken der Wirtskönigin angeklammert. Ihre zahlreiche Nachkommenschaft kopuliert im Mutternest (vgl. Anergates atratulus). Wie die Ausbreitung erfolgt, ist unbekannt.
Verwandtschaftlich leitet sich Teleutomyrmex aus der Gattung Tetramorium ab.
Ergänzung 30. Aug. 2007: Die arbeiterinlose sozialparasitische Ameise Teleutomyrmex schneideri wurde nun erstmals in Spanien nachgewiesen. Zwei Gynen fanden sich in Bodenfallen im Norden der Provinz Léon auf 1.660 m Meereshöhe. Das sind ungefähr 700 km westlich des nächstgelegenen Fundortes in den französischen Pyrenäen. Der Fund unterstreicht, wie außerordentlich verstreut und isoliert die Vorkommen dieser Art sind (Espadaler & Cuesta 2006).”

From AntWeb, on 14/03/2020, from 12:35 till 12:55:

“Genus: Teleutomyrmex Kutter, 1950.
Current Valid Name:
Tetramorium.
Teleutomyrmex Kutter, 1950a: 82. Type-species: Teleutomyrmex schneideri, by original designation.
Taxonomic history:
Teleutomyrmex in Myrmicinae, Tetramoriini: Kutter, 1950a: 81 [subtribe Teleutomyrmini].
Teleutomyrmex as junior synonym of Tetramorium: Ward et al., 2015 10.1111/syen.12090: 16.
Genus Teleutomyrmex references:
Bernard, 1967a: 239 (diagnosis); Bolton, 1976: 309 (diagnosis, review of genus); Bolton, 1994: 106 (synoptic classification); Bolton, 1995a: 1053 (census); Bolton, 1995b: 403 (catalogue); Sanetra & Buschinger, 2000: 95 (phylogeny).
Distribution:
Geographic regions: Not found on any curated Geolocale/Taxon lists.”

“Species: Tetramorium inquilinum Ward et al., 2015.
Tetramorium inquilinum Ward et al., 2015 10.1111/syen.12090: 16.
Taxonomic history:
Replacement name for Tetramorium schneideri: (Kutter). [Junior secondary homonym of Tetramorium schneideri: Ward et al., 2015 10.1111/syen.12090: 16.
Distribution:
Geographic regions (According to curated Geolocale/Taxon lists):
Europe: France, Switzerland, Ukraine.
Biogeographic regions (According to curated Bioregion/Taxon lists):
Palearctic.
Specimen Habitat Summary:
Elevations: collected at 2050 m.”

“Species: Tetramorium kutteri (Tinaut, 1990).
Teleutomyrmex kutteri Tinaut, 1990b: 202, figs. 1-3 (q.m.) SPAIN. Palearctic. HOL.
Taxonomic history:
Combination in Tetramorium: Ward et al., 2015 10.1111/syen.12090: 16.
Distribution:
Geographic regions (According to curated Geolocale/Taxon lists):
Europe: Spain.
Biogeographic regions (According to curated Bioregion/Taxon lists):
Palearctic.
Specimen Habitat Summary:
Found most commonly in these microhabitats: 1 times Juniperus-Genista brushwood.
Collected most commonly using these methods: 1 times Hand.
Elevations: collected from 1660 - 2250 meters, 1955 meters average.”

“Species: Tetramorium seiferti (Kiran & Karaman, 2017).
Teleutomyrmex seiferti Kiran & Karaman, 2017: 148, figs. 3a, 4a, 5a, 6-8 (q.m.) TURKEY. Palearctic. Primary type information: Holotype (gyne) from Turkey, Artvin, Yusufeli, 3 km NW of Kınalıçam Village, N 40° 45' 36", E 41° 34' 46", 1801 m above sea level (a.s.l.), 25.VI.2013, 13 / 1592c, leg. K. Kiran, C. Karaman & V. Aksoy (in Collection of Biological Department, Trakya University, Edirne, Turkey). Paratypes: 16 gynes, 7 males from same nest as the holotype (14 gynes, 5 males in Collection of Biological Department, Trakya University, Edirne; 1 gyne, 1 male in Sofia University, Bulgaria; 1 gyne, 1 male in Senckenberg Museum of Natural History Görlitz, Germany.
Taxonomic history:
[Note: Kiran et al., 2017 PDF: 146, retain the paraphyletic genus Teleutomyrmex.].
Distribution:
Geographic regions (According to curated Geolocale/Taxon lists):
Asia: Turkey.
Biogeographic regions (According to curated Bioregion/Taxon lists):
Palearctic.
Specimen Habitat Summary:
Elevations: collected at 1801 m.”

“Species: Tetramorium buschingeri (Lapeva-Gjonova, 2017).
Teleutomyrmex buschingeri Lapeva-Gjonova, 2017: 151, figs. 3b, 4b, 5b (q.) BULGARIA. Palearctic. Primary type information: Holotype (dealated gyne) from Bulgaria, Eastern Rhodopes Mt., Chernichino Village, N 41° 35' 29.71", E 25° 50' 55.03", 640 m a.s.l., 25.IV.2012, leg. A. Lapeva-Gjonova; deposited in the museum collection of Sofia University, Bulgaria (BFUS). Paratypes (1 dealated gyne) from the same nest as the holotype, deposited in the National Museum of Natural History in Sofia (NMNHS), Bulgaria.
Taxonomic history:
[Note: Kiran et al., 2017 PDF: 146, retain the paraphyletic genus Teleutomyrmex.].
Distribution:
Geographic regions (According to curated Geolocale/Taxon lists):
Europe: Bulgaria.
Biogeographic regions (According to curated Bioregion/Taxon lists):
Palearctic.”

From AntWiki, on 14/03/2020, from 15:35 till 16:20:

“Teleutomyrmex.
This genus is not in use as it is currently considered to be a junior synonym of Tetramorium.
Nomenclature.
The following information is derived from Barry Bolton's New General Catalogue, a catalogue of the world's ants.
TELEUTOMYRMEX [junior synonym of Tetramorium].
Teleutomyrmex Kutter, 1950: 82. Type-species: Teleutomyrmex schneideri, by original designation.
Teleutomyrmex junior synonym of Tetramorium: Ward et al., 2014: 15.”

“Teleutomyrmex schneideri.
This taxon is a junior homonym and has been replaced by Tetramorium inquilinum.
Nomenclature.
The following information is derived from Barry Bolton's New General Catalogue, a catalogue of the world's ants.
schneideri. Teleutomyrmex schneideri Kutter, 1950a: 82, figs. 1-23 (q.m.) SWITZERLAND.
[Junior secondary homonym of Tetramorium schneideri Emery, 1898c: 145.]
Combination in Tetramorium: Ward, et al. 2015: 76.
Status as species: Stumper, 1951: 129; Brun, 1952: 73; Gösswald, 1953: 81; Bernard, 1967: 240 (redescription); Bolton, 1976: 309 (redescription); Kutter, 1977c: 167; Dlussky, Soyunov & Zabelin, 1990: 210; Bolton, 1995b: 403; Casevitz-Weulersse & Galkowski, 2009: 494; Borowiec, L. 2014: 170; Kiran, et al. 2017: 146.
Replacement name: Tetramorium inquilinum Ward, et al. 2015: 76.”

“Tetramorium inquilinum.
Tetramorium inquilinum is a workerless inquiline within the nests of Tetramorium caespitum and Tetramorium impurum.
Distribution.
Distribution based on Regional Taxon Lists.
Palaearctic Region: France, Iberian Peninsula, Russian Federation, Spain, Switzerland (type locality), Turkmenistan.
Biology.
As reported by Hölldobler and Wilson (1990) - This remarkable species was discovered by Heinrich Kutter (1950a) at Saas-Fee, in an isolated valley of the Swiss Alps near Zermatt. Its behavior has been studied by Stumper (1950) and Kutter (1969), its neuroanatomy by Brun (1952), and its general anatomy and histology by Gösswald (1953). A second population has been reported from near Briançon in the French Alps by Collingwood (1956) (see also Wegnez et al. 2015), a third in the French Pyrenees by Buschinger (1987c), and still others in the Spanish Sierra Nevada by Tinaut Ranera (1981). The latter was described as a separate species, Tetramorium kutteri (as Teleutomyrmex kutteri), by Tinaut Ranera (1990). Appropriately, the name Teleutomyrmex means "final ant."
The populations of T. schneideri, like those of most workerless parasitic ant species (Wilson, 1963), are small and isolated. The Swiss population appears to be limited to the eastern slope of the Saas Valley, in Juniper-Arctostaphylos woodland ranging from 1,800 to 2,300 m in elevation. The ground is covered by thick leaf litter and sprinkled with rocks of various sizes, providing, in short, an ideal environment for ants. The ant fauna is of a typically boreal European complexion, comprising the following free-living species listed in the order of their abundance (Stumper, 1950): Formica fusca, F. lugubris, Tetramorium caespitum, Leptothorax acervorum, L. tuberum, Camponotus ligniperda, Myrmica lobicornis, M. sulcinodis, Camponotus herculeanus, Formica sanguinea, F. rufibarbis, F. pressilabris, and Manica rubida. For some unexplained reason this little assemblage is extremely prone to social parasitism. Formica sanguinea is a facultative slavemaking species, preying on the other species of Formica. Doronomyrmex (= Leptothorax) pacis, a workerless parasite living with Leptothorax acervorum, was discovered by Kutter as a genus new to science in the Saas-Fee forest in 1945. In addition, Kutter and Stumper found Epimyrma (= Myrmoxenus) stumperi in nests of Leptothorax tuberum, as well as two parasitic Leptothorax, goesswaldi and kutteri, in nests of L. acervorum (Kutter, 1969).
Tetramorium inquilinum is a parasite of Tetramorium caespitum and Tetramorium impurum. Like so many other social parasites, it is phylogenetically closer to its host than to any of the other members of the ant fauna to which it belongs. In fact, it may have been derived directly from a temporarily free-living offshoot of this species, since T. caespitum and T. impurum (the host species at Briançon and in the Pyrenees) are the only nonparasitic tetramoriines known to exist at the present time through most of central Europe. It is difficult to conceive of a stage of social parasitism more advanced than that actually reached by Tetramorium inquilinum. The species occurs only in the nests of its hosts. It lacks a worker caste, and the queens contribute in no visibly productive way to the economy of the host colonies. The queens are tiny compared with most ants, especially other tetramoriines; they average only about 2.5 mm in total length. They are unique among all known social insects in being ectoparasitic. In other words, they spend much of their time riding on the backs of their hosts (Figure 12-1). The T. inquilinum queens display several striking morphological features that are correlated with this peculiar habit. The ventral surface of the gaster (the large terminal part of the body) is strongly concave, permitting the parasites to press their bodies close to those of their hosts. The tarsal claws and arolia are unusually large, permitting the parasites to secure a strong grip on the smooth chitinous body surface of the hosts. The queens have a marked tendency to grasp objects. Given a choice, they will position themselves on the top of the body of the host queen, either on the thorax or the abdomen. Deprived of the nest queen, they will then seize a virgin Tetramorium queen, or a worker, or a pupa, or even a dead queen or worker. Stumper observed a case in which six to eight T. inquilinum queens simultaneously grasped one Tetramorium queen, completely immobilizing her. The mode of feeding of T. inquilinum is not known with certainty. The adults are evidently either fed by the host workers through direct regurgitation or else share in the liquid regurgitated to the host queen. In any case, they are almost completely inactive most of the time. The T. inquilinum adults, especially the older queens, are highly attractive to the host workers, who lick them frequently. According to Gösswald, large numbers of unicellular glands are located just under the cuticle of the thorax, pedicel, and abdomen of the queens; these are associated with glandular hairs and are believed to be the source of a special attractant for the host workers. The abdomens of older T. inquilinum queens become swollen with fat body and ovarioles, as is shown in Figure 12-1. This physogastry is made possible by the fact that the intersegmental membranes are thicker and more sclerotized than is usually the case in ant queens and can therefore be stretched more. Also, the abdominal sclerites themselves are widely overlapping in the virgin queen, so that the abdomen can be distended to an unusual degree before the sclerites are pulled apart. The ovarioles increase enormously in length, discard their initial orientation, and infiltrate the entire abdomen and even the postpetiolar cavity.
From one to several physogastric queens are found in each parasitized nest, usually riding on the back of the host queen. Each lays an average of one egg every thirty seconds. The infested Tetramorium colonies are typically smaller than uninfested ones, but they still contain up to several thousand workers. The Tetramorium queens also lay eggs, and these are capable of developing into either workers or sexual forms (Buschinger, personal communication). Consequently the brood of a parasitized colony consists typically of eggs, larvae, and pupae of T. inquilinum queens and males mixed with those of Tetramorium workers.
The bodies of the T. inquilinum queens bear the mark of extensive morphological degeneration correlated with their loss of social functions. The labial and postpharyngeal glands are reduced, and the maxillary and metapleural glands are completely absent. The mandibular glands, on the other hand, are apparently normal. In addition, the queens possess a tibial gland, the function of which is unknown. The integument is thin and less pigmented and sculptured in comparison with that of Tetramorium; as a result of these reductions the queens are shining brown, an appearance that contrasts with the opaque blackish brown of their hosts. The sting and poison apparatus are reduced; the mandibles are so degenerate that the parasites are probably unable to secure food on their own; the tibial-tarsal cleaning apparatus is underdeveloped; and, of even greater interest, the brain is reduced in size with visible degeneration in the associative centers. In the central nerve cord, ganglia 9-13 are fused into a single piece. The males are also degenerate. Their bodies, like those of the males of a few other extreme social parasites, are "pupoid," meaning that the cuticle is thin and depigmented, actually greyish in color; the petiole and postpetiole are thick and provided with broad articulating surfaces; and the abdomen is soft and deflected downward at the tip.
In its essentials the life cycle of Tetramorium inquilinum resembles that of other known extreme ant parasites. Mating takes place within the host nest. The fecundated queens then either shed their wings and join the small force of egg layers within the home nest or else fly out in search of new Tetramorium nests to infest. Stumper found that the queens could be transferred readily from one Tetramorium colony to another, provided the recipient colony originated from the Saas-Fee. However, Tetramorium colonies from Luxembourg were hostile to the little parasites. Less surprisingly, ant species from the Saas-Fee other than Tetramorium caespitum always rejected the T. inquilinum. However, Buschinger (personal communication) has pointed out that the Saas-Fee population could be caespitum or impurum, or a mixture of both. In other words, the transfer might have been attempted across species.
Nomenclature.
The following information is derived from Barry Bolton's New General Catalogue, a catalogue of the world's ants.
schneideri. Teleutomyrmex schneideri Kutter, 1950a: 82, figs. 1-23 (q.m.) SWITZERLAND.
[Junior secondary homonym of Tetramorium schneideri Emery, 1898c: 145.]
Combination in Tetramorium: Ward, et al. 2015: 76.
Status as species: Stumper, 1951: 129; Brun, 1952: 73; Gösswald, 1953: 81; Bernard, 1967: 240 (redescription); Bolton, 1976: 309 (redescription); Kutter, 1977c: 167; Dlussky, Soyunov & Zabelin, 1990: 210; Bolton, 1995b: 403; Casevitz-Weulersse & Galkowski, 2009: 494; Borowiec, L. 2014: 170; Kiran, et al. 2017: 146.
Replacement name: Tetramorium inquilinum Ward, et al. 2015: 76.
inquilinum. Tetramorium inquilinum Ward, Brady, Fisher & Schultz, 2014: 16.
Replacement name for schneideri Kutter, 1950a: 82. [Junior secondary homonym of Tetramorium schneideri Emery, 1898c: 145.]”

“Tetramorium kutteri.
This species is an inquiline. Queens live in the nest of a different ant species, have no workers and are entirely dependent on their hosts for food. The queens produce eggs that are cared for and raised to maturity by the host workers.
Distribution.
Distribution based on Regional Taxon Lists.
Palaearctic Region: Iberian Peninsula, Spain (type locality).
Nomenclature.
The following information is derived from Barry Bolton's New General Catalogue, a catalogue of the world's ants.
kutteri. Teleutomyrmex kutteri Tinaut, 1990b: 202, figs. 1-3, photos. 1-2 (q.m.) SPAIN. Combination in Tetramorium: Ward et al., 2014: 16.”

“Tetramorium seiferti.
A Turkish social parasite found in the nest of a of Tetramorium species nesting under a stone in a pine forest.
Identification.
Kiran, et al. (2017) - Gynes differ from Tetramorium schneideri by larger CS, HTL / CS, PPW / CS, and DLO / CS (Tab. 1). They differ from Tetramorium kutteri by larger CS and DLO / CS, and by much longer and more erect pilosity on appendages and whole body. They differ from Tetramorium buschingeri by smaller CW / CL, larger DLO / CS, by the dorsolateral margins of the propodeum forming distinct carinae, and by much less developed microsculpture on lateral mesosomal sclerites and petiole.
Distribution.
Distribution based on Regional Taxon Lists.
Palaearctic Region: Turkey (type locality).
Biology.
Kiran, et al. (2017) - The type series was collected from a Tetramorium cf. chefketi nest located under a stone in a Pinus sylvestris L. forest older than 100 years. There are many trees about to die because of their old age, and therefore there are very large sun exposed areas on the forest ground. The ground is not inclined near the nest because the nest site is placed on the top of the hill.
The habitat is similar to that reported by Tinaut (1990) for the type locality of Teleutomyrmex kutteri.
Nomenclature.
The following information is derived from Barry Bolton's New General Catalogue, a catalogue of the world's ants.
buschingeri. Teleutomyrmex seiferti Kiran & Karaman, in Kiran, et al. 2017: 148, figs. 3a, 4a, 5a, 6-8 (q.m.) TURKEY.
[Note: Kiran, et al. 2017: 146, retain the paraphyletic genus Teleutomyrmex.]
Unless otherwise noted the text for the remainder of this section is reported from the publication that includes the original description.
Description.
Queen.
Head in full face view not much wider than long (CL / CW 0.951 - 1.000), lateral sides convex and rounding to slightly concave posterior margin, anterior part of head narrower than posterior one. Eyes protruding and small (EL / CS 0.224 - 0.250), occupying less than one quarter of lateral head side, ocelli relatively well developed, distance of posterior ocelli rather large (DLO / CS 0.244 - 0.275), anterior clypeal margin deeply concave, posterior margin broadly convex, mandibles atrophied, largely triangular, and with pointed apex. Antennal scape as long as head length (SL / CL 0.974 - 1.000), slightly surpassing posterior margin of head, funiculus 10-segmented, 3rd funicular segment slightly longer than 4th one and almost twice as long as 2nd; three apical segments forming a small club.
Pronotum narrower than head (PNW / CS 0.732 - 0.835), anterolaterally with small angles, in lateral view scutum raised over pronotum, dorsal surface of scutum and prescutum straight, scutellum higher than scutum, metanotum very small, like a tooth. Dorsolateral margins of propodeum developed as distinct carinae, the longer dorsal and the shorter declivitous profile of carinae forming an obtuse angle with a blunt tooth at the position where spines normally are based; distance of propodeal spiracle from posterior margin of propodeum twice as long as the distance from anterior margin to spiracle. Petiolar scale in profile triangular, with a nearly linear frontal and broadly convex caudal profile. Dorsal outline of postpetiole convex in lateral view. Gaster typically dorso-ventrally flattened in virgin females.
Head smooth and shiny, only genae between antennal sockets and compound eyes microreticulate. Mesosoma and petiole microalveolate. In dorsal view, postpetiole superficially microreticulate and gaster glabrous.
Head and hind tibiae with long erect hairs, scape and hind femur with dense semi-erect hairs. Dorsum of mesosoma, petiole and postpetiole with brush-like hairs, anepisternum and gaster almost bare, katepisternum with long decumbent hairs.
Clypeus, mandibles, antennae, and legs yellow. Rest of head, mesosoma, and petiole brown; postpetiole brownish yellow; gaster brownish yellow except yellow lateral sides and articulations.
Male.
In full face view head wider than long (CL / CW 0.910 - 0.935), lateral sides convex, rounding to straight posterior margin, anterior part of head slightly narrower than posterior one, eyes relatively big (EL / CS 0.233 - 0.272), located in the middle of head sides, ocelli well developed, distance of posterior ocelli rather large (DLO / CS 0.247 - 0.282), anterior margin of clypeus slightly notched medially, posterior margin broadly convex, mandibles atrophied and very small. Antennal scape short (SL / CS 0.871 - 0.907), scape hardly reaching posterior margin of head, funiculus 10-segmented, last three articles form a club.
Pronotum short and distinctly narrower than head (PNW / CS 0.762 - 0.805). With mesosoma in lateral view, mesonotum not raised as in female, its dorsum almost straight; scutellum strongly raised, metanotum lower than scutellum; dorsal surface of propodeum sloping posteriorly, slightly longer than declivitous surface and joining by an obtuse angle, and forming here a small tooth. Propodeal spiracle located close to anterior propodeal border. Dorsal part of petiole in profile almost rectangular, petiole distinctly wider than the distance between propodeal teeth, postpetiole very wide and short in dorsal view. Gaster dorsoventrally flattened.
Posterior margin of subgenital plate convex. Shape of sagitta and subgenital plate more similar to Teleutomyrmex kutteri on the contrary volsella completely different from T. kutteri and T. schneideri.
Head densely microreticulate and matt, mesosoma and petiole densely microalveolate. Postpetiole, first and second abdominal tergites with very dilute microreticulum and rest of gaster glabrous.
Long hairs absent or very rare. Short, erect and suberect hairs present on head, scape, mesosoma, petiole, and postpetiole. Gaster bare.
Lower parts of head, antennae, and legs yellow; rest of head, mesosoma, petiole, and postpetiole yellowish brown.
Gaster brownish yellow.
Type Material.
Holotype (gyne) from Turkey, Artvin, Yusufeli, 3 km NW of Kınalıçam Village, N 40° 45' 36", E 41° 34' 46", 1801 m above sea level (a.s.l.), 25.VI.2013, 13 / 1592c, leg. K. Kiran, C. Karaman & V. Aksoy (in Collection of Biological Department, Trakya University, Edirne, Turkey). Paratypes: 16 gynes, 7 males from same nest as the holotype (14 gynes, 5 males in Collection of Biological Department, Trakya University, Edirne; 1 gyne, 1 male in Sofia University, Bulgaria; 1 gyne, 1 male in Senckenberg Museum of Natural History Görlitz, Germany.
Etymology.
The species is dedicated to Dr. Bernhard Seifert for his great contributions to this study.”

“Tetramorium buschingeri.
A Bulgarian social parasite found in the nest of a of Tetramorium species nesting under a rock in a grassland.
Identification.
Kiran, et al. (2017) - Gynes differ from the other three Teleutomyrmex species by the absence of any carinae or teeth on dorsal surface of propodeum, by a very short dorsal profile of propodeum, by a much stronger developed reticulate or alveolate microsculpture covering the whole surface of lateral mesosoma and petiole and by smaller CL / CW. Furthermore, they differ from Tetramorium seiferti by a much smaller DLO / DFC (0.567 - 0.600 vs. 0.667 - 0.786). Additional differences to Tetramorium schneideri are a larger HTL / CS and a smaller EL / CS, and to Tetramorium kutteri a much smaller CL / CW, larger PW / CS and HTL / CS, and smaller ClyW / CS.
Distribution.
Distribution based on Regional Taxon Lists.
Palaearctic Region: Bulgaria (type locality).
Biology.
Kiran, et al. (2017) - Host: The two dealated queens were found on the back of a dealated queen of Tetramorium cf. chefketi together with few host workers in the nest. The host ant species belongs to the T. chefketi species complex in contrast to the hosts from the T. caespitum / impurum complex reported for Tetramorium schneideri and Tetramorium kutteri in Europe.
The nest of Tetramorium cf. chefketi with Teleutomyrmex buschingeri was found under a stone on a southern slope of a dry grassland situated in an oak forest. The habitat type is quite different from the known habitats of related species. The altitude of 640 m, where the species was sampled, is notably less than the altitudes of the localities of the other ultimate ant parasites (1600 - 2300 m) (excluding one in Turkmenistan).
Nomenclature.
The following information is derived from Barry Bolton's New General Catalogue, a catalogue of the world's ants.
buschingeri. Teleutomyrmex buschingeri Lapeva-Gjonova, in Kiran, et al. 2017: 151, figs. 3b, 4b, 5b (q.) BULGARIA.
[Note: Kiran, et al. 2017: 146, retain the paraphyletic genus Teleutomyrmex.]
Unless otherwise noted the text for the remainder of this section is reported from the publication that includes the original description.
Description.
Queen.
Head in full face view wider than long (CL / CW 0.904), its lateral sides distinctly convex, anterior part of head narrower than posterior one, eyes small (EL / CS 0.228), lateral ocelli well developed, lateral ocelli less distant from each other than in Teleutomyrmex seiferti (DLO / CS 0.219). Anterior clypeal margin deeply concave and posterior margin rather convex. Mandibles reduced and triangular, with a pointed apex. Antennal scape distinctly longer than head length, exceeding the dorsal margin of head. Funiculus 10-segmented, 2nd segment 0.043 mm, 3rd segment 0.067 mm, 4th segment 0.055 mm long, the three apical segments form a small club. Pronotum narrower than head (PNW / CS 0.747), anterolaterally with small angles. Mesonotum raised over pronotum, its dorsum straight, scutellum higher than meso notum. Dorsal surface of propodeum much shorter than declivitous surface, both surfaces form a convex outline without any traces of propodeal teeth. Propodeal spiracle located near to anterior border of propodeum. Petiolar scale in profile triangular with straight frontal and convex posterior surface. Dorsal outline of postpetiole convex in lateral view. Gaster in virgin gynes typically dorso-ventrally flattened. Hind tibiae very long (HTL / CS 0.994) distinctly longer than in Teleutomyrmex kutteri and T. schneideri.
Clypeus and frons up to the level of anterior ocellus smooth and shiny, between compound eye and antennal socket densely microreticulate. Mesosoma and petiole densely microalveolate and matt, postpetiole slightly microreticulate and more shiny. Gaster glabrous.
Head with moderately long erect hairs. Scape with semi-erect hairs. Mesosoma and petiole with brush like hairs. Postpetiole with very short brush like hairs. Gaster bare. Hind femora with long subdecumbent hairs. Tibiae with very long, erect hairs. Katepisternum and lateral portion of propodeum with very few and short decumbent hairs.
Clypeus, mandibles, antennae, and legs yellow. Head, mesosoma and petiole dark brown. Postpetiole yellowish brown, first gaster segment brown, the rest brownish yellow.
Type Material.
Holotype (dealated gyne) from Bulgaria, Eastern Rhodopes Mt., Chernichino Village, N 41° 35' 29.71", E 25° 50' 55.03", 640 m a.s.l., 25.IV.2012, leg. A. Lapeva-Gjonova; deposited in the museum collection of Sofia University, Bulgaria (BFUS). Paratypes (1 dealated gyne) from the same nest as the holotype, deposited in the National Museum of Natural History in Sofia (NMNHS), Bulgaria.
Etymology.
The species is dedicated to Prof. Alfred Buschinger who has made great contributions to the study of socially parasitic ants.”

“Key to parasitic Tetramorium species.
This queen key is based on: Kiran, K., Karaman, C., Lapeva-Gjonova, A. & Aksoy, V. 2017. Two new species of the “ultimate” parasitic ant genus Teleutomyrmex Kutter, 1950 from the Western Palaearctic. Myrmecological News 25: 145-155.
Originally assigned their own genus, Teleutomyrmex, these ants prey on other Tetramorium species. The species Tetramorium atratulum is not included in this key and bears no resemblance to the other parasites in the genus.
Males of T. buschingeri have yet to be collected.
1
Gynes . . . . . 2
Males . . . . . 5
2
Carinae or teeth on dorsal surface of propodeum absent, dorsal profile of propodeum much shorter than the declivitous one. All lateral surfaces of mesosoma and petiole covered by a well-developed reticulate or alveolate microsculpture. Head length index CL / CW < 0.945. Southern Balkans . . . . . Tetramorium buschingeri.
Carinae or teeth on dorsal surface of propodeum present, dorsal profile of propodeum not much shorter than the declivitous one. Surfaces of lateral mesosoma and petiole only in patches covered by a reticulate or alveolate microsculpture or completely smooth. Head length index CL / CW > 0.945 . . . . . 3
3
Scape long, SL / CS > 1.00. Distance of frontal carinae clearly larger than petiolar width, DFC / PW > 1.096. Size small, CW < 464 μm. Scapes and tibiae with weaker, largely decumbent pilosity. Southern Iberia . . . . . Tetramorium kutteri.
Scape shorter, SL / CS < 1.00. Distance of frontal carinae not much larger than petiolar width, DFC/PW < 1.096. Size larger, CW > 464 μm. Scapes and tibiae with profuse erect or suberect pilosity . . . . . 4
4
Ratio of distance between lateral ocelli and large diameter of complex eye larger: DLO / EL 0.93 - 1.11. Katepisternum with many long decumbent hairs, posterior corners of head posterior of the eyes smooth, absolute scape length larger: SL > 457 μm. Anatolia . . . . . Tetramorium seiferti.
Ratio of distance between lateral ocelli and large diameter of complex eye smaller: DLO / EL 0.70 - 0.80. Katepisternum without or only with a few decumbent hairs, posterior corners of head posterior of the eyes densely microreticulate, absolute scape length smaller: SL < 457 μm. Alps and Pyrenees . . . . . Tetramorium inquilinum.
5
Anterior clypeal margin straight . . . . . 6
Anterior clypeal margin concave medially . . . . . Tetramorium seiferti.
6
Subgenital plate broadly convex, sagitta with sinusoidal shape . . . . . Tetramorium kutteri.
Subgenital plate slightly concave, sagitta broadly convex . . . . . Tetramorium inquilinum.”

And the species not included in all 3 pages:

The undescribed species from Turkmenistan… See Dlussky, et al. 1990.
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Mittwoch 25. März 2020, 19:49

Teleutotje hat geschrieben:Maybe this belongs here but... Personally I think this is a workerless inquiline, NOT extreme...

http://antcat.org/documents/6525/messer ... enopte.pdf


And now two more species:

https://zookeys.pensoft.net/article/469 ... pdf/393900
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Mittwoch 1. April 2020, 21:12

Teleutotje hat geschrieben:Now, a new contribution to Myrmecological News. I'm still reading it but it looks great.

de la Mora, A., Sankovitz, M. & Purcell, J., 2020, "Ants (Hymenoptera: Formicidae) as host and intruder: recent advances and future directions in the study of exploitative strategies." Myrmecological News, Vol. 30, 53-71.

https://myrmecologicalnews.org/cms/inde ... Itemid=435
or
https://myrmecologicalnews.org/cms/inde ... format=raw and https://myrmecologicalnews.org/cms/inde ... format=raw


And a review of it: https://blog.myrmecologicalnews.org/202 ... -intruder/ .
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Sonntag 5. April 2020, 11:13

Teleutotje hat geschrieben:
Teleutotje hat geschrieben:Maybe this belongs here but... Personally I think this is a workerless inquiline, NOT extreme...

http://antcat.org/documents/6525/messer ... enopte.pdf


And now two more species:

https://zookeys.pensoft.net/article/469 ... pdf/393900


Prof. Buschinger: viewtopic.php?f=23&t=367&start=240#p20355
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Re: The extreme, workerless inquilines.

Beitragvon Teleutotje » Freitag 29. Mai 2020, 22:24

Tinaut, A., Martínez-Ibáñez, M. D., Ruano, F., 2007, “Inventário de las especies de formícidos de Sierra Nevada, Granada (España) (Hymenoptera, Formicidae).” or “Inventory of the ant species of Sierra Nevada, Granada (Spain) (Hymenoptera, Formicidae).” Zoologica Baetica, vol. 18, p. 49-68.
- 20 pp., 0 figs. - [2007-??-??].

93 references.
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